You might consider the Tamron or Sigma 70-200mm f/2.8. I know it's a bit out of your listed budget, but the Tamron can be found for $650-700 if you shop around. The Sigma is a bit more. Either will have better optics, and a faster and constant aperture across all focal lengths and both are macro lenses.
I looked at both, as well as the Canon 70-200mm f/2.8L non-IS. I initially gave the Tamron a shot. IIRC, the Tamron is sharper than the Sigma and I was quite happy with the optics. People say it is slow to focus, but as with my Tamron 17-50mm f/2.8, I think the noise of the motor is misleading. I haven't found the Canon to be appreciably faster at focusing, certainly not $500 faster. The Tamron is also significantly lighter and has a 6 year warranty.
The Tamron has a nice auto/manual focus collar that you simply have to shift forward and back and worked fine for me, which is a reasonable compromise over having full-time-manual. You might lose a little focus lock when shifting the collar, if you're into auto focusing and then manually tweaking.
3rd party lenses aren't for everyone though, but I believe they're fine for the hobbyist on a budget. I'm not sure how well Tamron's play with Nikon's, but I love my Tamron 17-50mm f/2.8 - that's vastly better than the kit lenses. I ended up with the Canon version, mostly because I didn't want to second guess my purchase when I had the cash for the Canon and I wanted the full-time-manual that the USM motor provides. I gave up macro by doing so, which I do miss. The min focus distance for the Canon is a whopping 3 feet!
Just something to consider. It's pricey, and no IS/VR, but very sharp and faster (the f/2.8 makes up for the lack of IS/VR somewhat.) FWIW, I use my 70-200mm a lot more than I originally expected.