They're variable (that's why the Kinesis Advantage has middle finger column sitting deeper than the rest) but they're not staggered. The zig-zag staggered layout I'm talking about was devised to fit the typewriter pushrods. We don't need it anymore.
This is not natural movement for the hands or fingers, and no matter how fancy you make the switches and charge $300-400 or more, if you don't deal with the fundamental flaw of traditional keyboards, you have a mediocre keyboard. Even if the label on it says Realforce.
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I know the internet, and GeekHack especially are havens for argumentative people. Usually I keep my thoughts to myself but I've done a lot of research on desk and keyboard ergonomics during my undergraduate studies.
As a RN with a Bachelor's of Science in Kinesiology I have to say I can't agree with this. The staggered keyboard favors the medial angles of your index fingers. A straight keyboard would actually cause the shoulders to anatomically protract.
The biggest ergonomic factor is how far apart your hands are and the angle your keyboard is split at. If your keyboard is angled (studies show 5-12 degrees yields less hand fatigue) your ring finger and pinky finger will be aligned in a position that yields less force and stress on your digital muscles. I'll find some articles if you guys are really interested.
Edit: Also the trulyergonomic picture is seriously flawed. Anyone can test this by placing their hands on a regular keyboard and can see that they don't position their hands as shown in the picture. While the trulyergonomic keyboard favors proper body positioning more than regular keyboards their product is no more advantageous (layout wise) than using a Microsoft keyboard.
Switch-wise, the trulyergonomic keyboard would have an advantage over average ergonomic keyboards if it were to have tactile switches. PM me if you want an explanation on this.. it's kinda lengthy.